TRIBE CABERINI
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Astygisa Walker Gen. rev.

Type species: larentiata Walker (= vexillaria Guenée).

Synonyms: Alana Walker (type species rubiginata Walker = vexillaria); Apopetelia Wehrli (type species morosa Butler, Japan) syn. n.

The genus includes a number of dull reddish or maroon species, more robustly built than Petelia. Features of wing pattern include development of a strong white or yellow discal mark to the hindwing, zones of speckled white clouding on both wings, and, on the underside, relative uniformity, the border slightly and diffusely darker, with no dark discal spots.

The definitive feature is in the male abdomen where coremata of varying size occur between some or all segments from the apex to segment 4. In taxa with the coremata highly developed, they are supported by narrow bands of sclerotisation.

The male genitalia are more robust and heavily sclerotised than in Petelia, usually with strong setae apically and sometimes costally on the valves. The socii are more developed than in Petelia. The aedeagus vesica bears one or more strong cornuti that may themselves bear small subsidiary spines towards the apex (e.g. morosa and circularia Swinhoe).

In the female genitalia (Bornean species examined) the ballooning to the membrane dorsally just anterior to the ovipositor lobes is shared with Petelia. Diagnostic features are: the presence of a slight to moderate appendix bursae arising from near the base of the ductus and giving rise to the ductus seminalis; displacement of the ostium anteriorly onto segment 7 with development of a strong, often posteriorly serrate lamella vaginalis; strong fluting and sclerotisation of the ductus bursae, the ridges of the flutings bearing small spines; lack of a signum in the bursa (general scobination sometimes present).

The genus is Oriental, with its greatest morphological diversity seen from the Indian Subregion to Sundaland. In Sulawesi there is a complex of at least nine species with asymmetric structure of the uncus and valves; the names theclaria Walker and furva Warren are applicable. The complex probably also extends to the Lesser Sundas, as one specimen from Timor (slide 12897) has been located. The complex may bear a sister-relationship to vexillaria, indicated particularly by the sharing of a spined dorsal margin to the sacculus. Holloway (1991) presented an analysis of this group.

No life histories are available for the Bornean species, but Sato (1976) described the larva of two Japanese species. They are deep greenish yellow with the posterior margin of each abdominal segment pale yellow and the venter pale greenish yellow. The legs of T1 and the prolegs are tinged reddish purple. The spiracles are orange with a black rim. There is also a reddish purple form and intermediate forms. The host-plant was Hovenia (Rhamnaceae).

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