TRIBE CASSYMINI
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Heterostegane subtessellata Walker
   
Macaria subtessellata Walker, 1862, List Specimens lepid. Insects Colln Br. Mus., 26: 1648.
   
Terpnomicta quadrilineata Snellen, 1886, Tijdschr. Ent., 29: 49.


Heterostegane subtessellata


Diagnosis.
This and the next species are of similar size but can be distinguished by the generally darker appearance of contessellata, and the greater breadth of its forewing medial. The forewing postmedial in subtessellata is curved unevenly, concave basad, but more straight in contessellata except where angled basad at the costa and distad at the dorsum. In subtessellata there is a distinctive dark shading between medial and postmedial at the hindwing costa. The underside in contessellata is much more strongly and broadly fasciated. In the male genitalia the ventral portion in contessellata is small, rounded (rather than prominent, quadrate) and appears more contiguous with the dorsal arm.

Taxonomic notes. Sundanian specimens tend to be more densely speckled with red compared with Indian ones, so they appear somewhat darker. The hindwing of the submarginal at the forewing dorsum and at the position of M2 on the hindwing is less, and the veins between the postmedial and submarginal are not so strongly delineated with red. In the male genitalia, Bornean and Sumatran specimens have the distal part of the ventral portion of the valve narrower and more curved out at right angles. A syntype of quadrilineata (colour photograph examined) resembles typical Indian material more closely.

Geographical range. India, Nepal, Burma, Sundaland.

Habitat preference. The four Bornean specimens are all from lowland dipterocarp forest on various substrates, but not heath forest.

Biology. The larva has been described by Singh (1953) and Bell (MS). It is fusiform in shape, appearing flattened because of lateral flanges running the length of the body. Transverse folds across the abdominal segments give each segment a bluntly saw-like appearance in lateral view. The colour is pale green with thickish white subdorsal lines; the edge of the flange on each side is broadly whitish yellow and this colour continues over T1. The posterior margins of T3 to A7 are broadly transversely white above yellow below. The apex of the anal shield is pink.

The larva, when at rest, lies stretched full length along the midrib of leaflets of a compound leaf of the host-plant. Pupation is in a loose cell of silk incorporating leaflets and detritus. The moth sits with the wings semi-erect rather than flattened against the substrate.

The host-plants recorded have been species of Acacia.

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