SUBFAMILY ACRONICTINAE
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Craniophora malesiae sp. n.
Craniophora fasciata Moore sensu Holloway, 1976: 13.


Craniophora malesiae


Craniophora malesiae


20-21 mm, 21-22 mm. This species is slight larger than fasciata Moore ( = divisa Moore and nigrostriata Pagenstecher) from the Indian subregion but also identified from Java, Bali, Kei, Ambon and the Bismarck group (Squally I.). It has a greater degree of sexual dimorphism in the hind- wing, that of the male being more triangular, a warm bone colour, uniform except for a very diffuse medial band and, in unworn specimens, strong dark lunules on the fringes. The primary distinction is in the male genitalia: cornuti in the distal row (on a lateral lobe) are three to five in number, long, separate; there are also two separate ones basally, one triangular, one digitate. In fasciata only the distal group is present, the cornuti more numerous, fused into a comb. Two other species occur in this fasciata complex: one from New Guinea with an elongate vesica, the distal row of cornuti long, the cornuti very short, not on a lateral lobe with, just basal to it, a coarsely scobinate lobe another lobe bearing two short, conical cornuti; one from Queensland (slide 14021) and New Caledonia (Holloway, 1979: fig.98,as fasciata) with two short rows of small cornuti set centrally and at three quarters on the vesica with the vesica distal to the latter coarsely scobinate.

Holotype  BRUNEI: 300m, Ulu Temburong rainforest, 30.6.79 (Lt. Col. G. Allen) BM noctuid slide 13961.

Paratypes: 2 BRUNEI: 1618 m Bukit Retak, Montane forest (Lt. Col M.G. Allen) BM noctuid slide 14041.

Geographical range.
N.E. Himalaya, Borneo, Java (and probably throughout Sundaland), Sulawesi, New Guinea, Bismarck Is.

Habitat preference.
Most Bornean material has been taken in upper montane forest. One specimen was taken in lowland dipterocarp forest.

Biology.
The larva of this species or fasciata was illustrated by Sugi (1987). It is bright pale green, with long black primary setae set on black pinacula; the black spots paired dorsolaterally on A8 are larger than the rest, just enclosed by converging yellow stripes that extend forwards from the claspers to anteriorly on A8. The thorax tapers to the head that is laterally black, frontally reticulated green.

Recorded hosts (Miyata, 1983; Sugi, 1987; Bell, MS) for fasciata (or possibly malesiae) are Olea, Ligustrum and Osmanthus (Oleaceae).

Bell (MS) described a larva similar to that illustrated by Sugi. The larva on the underside of young leaves of the host-plant and is generally sluggish, resting with the head turned round on one side. Pupation is in an ovoid cell in rotten wood, lined with silk that, with chewed wood particles, also blocks the opening to the cell.

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