Cyclodes Guenée
Type species: omma Hoeven, Java.
Synonym: Beregra Walker (type species replenens Walker, Sri Lanka, Bangladesh = omma).
The facies of
this genus is unmistakable, pale mauve and brown, particularly the antemedial
ocellus on the forewing. The antennae are very long, extending to beyond three‑quarters
of the length of the forewing, and are smooth, filiform in the male. The
clypeofrons is sparsely scaled, the scales tending to converge at a central
point. The labial palps are catocaline in proportion and orientation, albeit
with a somewhat short third segment. The foretibia in both sexes is modified
somewhat with hair tufts.
In the male
abdomen, the eighth tergite is anteriorly narrow, with a strong pair of
apodemes, and broadens to twice the anterior width distally. The sternite has
an ovate anterior lacuna surrounded by a slight frame with very shallow
apodemes on the anterior margin, but no lateral rods. This segment may be a
weakly developed form of the framed corematous type. There are no diaphragmal
lobes between the first and second tergites in both sexes. The genitalia have
the tegumen broadened each side, with a zone of hairs that may represent a
peniculus. The articulation of the tegumen and vinculum involves a marked
overlap that may include an paratergal sclerite, though the continuity of this exterior
portion of the overlap with the vinculum is hard to interpret. The valves are
slightly upcurved, simple, expanded subapically, with a small conical harpe at
the apex of the sacculus. The juxta is a transversely diamond‑shaped
plate. The aedeagus vesica is a simple tube, slightly swollen near the base.
In the female
genitalia, the ovipositor lobes are flattened, resembling the condition in
Limacodidae, a family also with some propensity for palms as host plants. The
ostium is situated between the seventh and eighth segments, the former with the
sternite only slightly smaller than the tergite. The ostium is half as wide as
the eighth segment. The ductus narrows abruptly from it and is mostly slender,
unsclerotised, and one third of the length of the corpus bursae. This latter is
an elongate ovate, scobinate throughout, but with the scobination stronger
subbasally and apically.
The scaled
clypeofrons, the structure of the basal tergites of the abdomen, and the
peniculus, paratergal area and valve structure of the male genitalia suggest
that the genus may be better placed as a basal member of the trifine sequence
of groups rather than with the catocalines. The sinuous hindwing fasciation is
reminiscent of that in genera such as Ortopla Walker.
The larva of the
type species is described below. The arrangement of the setation anteriorly
resembles somewhat that described for Donda sailendra Kobes on p. 17.
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