Arcte Kollar
Type species: polygrapha Kollar, India.
Synonym: Cocytodes Guenée (type species coerula Guenée, East Indies).
This genus has
numerous unique features, but is one of those that includes species with both
spined (coerula) and unspined (the rest) midtibiae, leading them to have
been split in the past between the Catocalinae and Ophiderinae (Holloway, 2005,
and references therein). The species are all large, robust. Males have a
massive corema in the eighth abdominal segment; the abdomen is also
distinguished by a scaleless, laminate plate dorsally that extends over several
of the more distal segments; it may be sound-producing. The two lobes in the
diaphragma between first and second tergites are shallow, triangular. The
facies is uniform through the genus as illustrated and as described below, the
rather ligneous forewing pattern contrasting with the iridescent blue banding
of the hindwing. The male antennae are filiform, smooth, and the hindwing
dorsum of that sex has extensive hair‑scales. The labial palps, though
upturned, have a very small third segment. The clypeofrons is scaled, the
scales converging on the axis and directed obliquely upwards at 45°; the axis
is unscaled but covered by this convergence.
Features of the
male abdomen have been mentioned above. The genitalia have a robust uncus with
an apical hook. The tegumen is rather swollen on each side. The valves are
simple, tongue like, upcurved, sometimes with small processes on the interior
margin of the sacculus. The juxta is a simple plate. The aedeagus is short,
straight, the vesica extending from it for half the length with a few small
patches of spines of different sizes and density, and also usually a spine or
other sclerotised process subbasally.
The female of the
type species has the ostium just posterior to the almost complete ring of the
eighth segment, with an extensive intersegmental membrane between it and the
seventh segment. The ductus is moderate, slender, unsclerotised but fluted. The
corpus bursae is an elongate pyriform, lightly corrugate and rugose to
scobinate throughout. The ductus seminalis arises from close to the base of
this.
The genus
contains four species and extends through the Indo‑Australian tropics to
Australia and Samoa.
The larvae are
striking and appear to feed exclusively on Urticaceae as discussed below.
The African genus
Pseudoarcte Viette has similar male antennae, forewing facies and
scaling of the clypeofrons, and may therefore be related, as may the
Neotropical Pararcte Hampson for the same reasons.
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