Belciana kala Prout
comb. rev. (Plate 1, Figs 25, 26)
Belciana
kala Prout, 1924, Bull. Hill
Mus. Witley, 1: 442.
Belciana kala Prout; Holloway,
1976: 32.
Donda kala Prout; Kobes, 1992, Heterocera
Sumatrana, 2: 85.
Belciana kala nepalensis Haruta, 1993, Tinea,
13 (Suppl. 3): 63.
Diagnosis. This is the largest Bornean species. It
has fine, dense, black, zig‑zag fasciation over the central part of the
forewing. The marginal zone with brown blocks has a row of ovate white spots
close to the actual margin. The hindwing is pale yellow over the basal half
with a dull brown‑grey border, more as in typical Donda.
Taxonomic note. Kobes (1992) transferred the species to Donda
on the basis of some hairs within the ommatidia of the eyes and other features,
but the male genitalia on balance indicate a closer relationship to Belciana.
The male abdomen has the eighth segment as in typical Belciana species,
but the valves of the genitalia are narrower, splayed away from the tegumen and
bearing a dense covering of dorsally directed hair setae as in B. striatovirens Moore (see below). The spine to the inner margin of
the sacculus is as in typical species, but there is an additional process more
basally. The distal spine is on a narrow detached section in ssp. nepalensis,
and the basal process is more acute. The aedeagus has an apical thorn, and the
vesica is more elongate and lacks the small curved sclerite of typical species.
The tegumen lacks a peniculus. The female genitalia follow a similar plan to
those of Donda and typical Belciana except the ostium is
broadened, lobed, funnel-like, the ventral zone of the corpus bursae is only
sclerotised and corrugated over a short distance, giving way immediately to the
distal zone which is elongate, slightly tapering.
Geographical range. Sundaland; Himalaya (ssp. nepalensis).
Habitat preference. The species is infrequent but ranges from
the lowlands to 1930m, mostly in forested localities. It may be slightly more
frequent over the middle of this range.
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