TRIBE CALPINI
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Plusiodonta Guenée

Type species: chalsytoides Guenée, Java, = coelonota Kollar, Indian Subregion.

Synonyms:
Deva Walker (type species conducens Walker, Sri Lanka = coelonota); Gadera Walker (type species incitans Walker, type locality not known [Mexico, Costa Rica]); Odontina Guenée (type species excavata Guenée, Réunion I.) praeocc.; Tafalla Walker (type species clavifera Walker, Honduras); Tinnodoa Nye (replacement name for Odontina).

Plusiodonta was also briefly discussed by Bänziger (1983). The forewing shape is more as in Oraesia than Calyptra, the pattern with golden or silvery areas on a dark ground. Oblique fasciation is more obscure but runs from the apex of the subbasal of the dorsum to the apex of the wing, i.e. more longitudinally oblique than in Calyptra or Oraesia. The male antennae are smooth or sparsely ciliate. The labial palps have the second segment deeply scaled, but the third is much more slender, slightly upturned.

In the male abdomen, the eighth segment is of the framed corematous type, the sternite modified to become rather shallow, with the corematous area almost obliterated, and with the lateral bands rather pinched in on each side. The uncus, scaphium and juxta resemble those of
Oraesia. The valves are very broad, particularly so distally, and the distal margin may be angular. As in Calyptra, there are processes from the inner margin of the sacculus, one more or less centrally and the other towards or at the valve apex (or there may be a sequence of three), with two in the apical zone around the margin). The aedeagus vesica is somewhat tubular, simple, and may have a cluster of spines distally.

The female has the ostium associated with the anterior of a rather elongated eighth segment, somewhat recessed within the seventh, where the sternite is distinctly shortened and distally rounded. The apodemes of the eighth segment are often reduced or vestigial. The ductus and bursa together are rather elongate and slender, sometimes with zones of scobination or sclerotisation. The ductus seminalis arises in various positions, sometimes (but not always) at the position of broadening of the corpus bursae.

The larva of
coelonota has been described by Moore (1884-1887), Sevastopulo (1946), Gardner (1947) and Bell (MS). It lacks prolegs on both A3 and A4, and A8 is slightly tumid dorsally. The head is orange, sometimes more reddish or reddish brown. The body is violet black to black, with velvety black patches and marked with grey patches, speckles, streaks and oblique bands, and sparsely spotted with blue. The spiracles are yellow. Sevastopulo (1946) noted a lateral white patch on A1 and A8, and dorsal white reticulation on A2. A4 is dorsally grey, the rest white. It is possible that more than one species is involved in these descriptions. The larva illustrated by Mutuura et al. (1965) is a very pale grey, marked with black dots and white patches; the head is orange.

The cocoon is described as spindle- or shuttle-shaped, made of white silk, covered with fragments of chewed leaf or bark, leading to a general grey, greenish or purplish-grey appearance. The cocoon is fixed along a twig or similar structure. The pupa lacks a bloom.

Most host records for this and other Oriental species (Miyata, 1983; Sugi, 1987; Robinson
et al., 2001) are from the Menispermaceae (Cocculus, Cyclea, Diploclisia and Stephania), but the larva has also been reared on Garcinia (Guttiferae) and Lepisanthes (Sapindaceae).

The genus is pantropical and extends into temperate latitudes. In the Indo-Australian tropics it extends east to Fiji and New Caledonia, both of which support endemic species (Robinson, 1975; Holloway, 1979).

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