TRIBE SCOLIOPTERYGINI
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Cosmophila Boisduval Gen. rev.

Type species: xanthindyma Boisduval, Mauritius, Réunion = flava Fabricius.

Synonyms:
Deinopalpus Holland (type species africana Holland, West Africa = leona Schaus); Deremma Walker (type species simulatrix Walker, Sierra Leone).

Definition of this genus has been provided under
Anomis on p. 221, with the caveat that some of the species tentatively assigned to it may prove to be better placed in Rusicada where the African fauna and the distribution of characters amongst the species has been more extensively surveyed than has been possible here. The species guttanivis Walker may be a case in point, as discussed below.

The core group of
Cosmophila was reviewed by Tams (1924a) and is essentially defined by: the contrasting yellow and grey of the forewing facies (also in scitipennis Walker); sclerotised processes from the interior of the valves; a well developed and complex sterigma around the ostium of the female. The bursa contains a signum consisting of a single curved spine, but a similar feature is present in Molopa (doubled), Gonitis and Lineopalpa horsfieldi Guenée, so the polarity of this character is unclear. The larvae of two of the species discussed by Tams (1924a) have appendiculate crochets, an unusual feature outside the noctuid subfamily Bagisarinae (e.g. as discussed by Holloway (1998)), though seen also in Marcillada Walker (see p. 206). They also differ in having the prolegs on A3 absent rather than reduced.

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