SUBFAMILY ARIOLICINI
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Labanda Walker

Type species: herbealis Walker, Sri Lanka.

Synonyms:
Bariana Walker (type species submuscosa Walker, Java); Gerbatha Walker (type species laticincta Walker = semipars Walker, Sri Lanka); Lazanda Walker (type species fasciata Walker, India); Pseudalea Turner (type species macrogastris Turner = huntei Warren, Queensland).

This and the next three genera have features of build and male genitalia in common as discussed on pp. 140-141.

The build is delicate, the forewings rather narrow, the abdomen, particularly in males, extending well beyond the hindwings in a spread specimen. The male antennae are usually filiform, but two species have them bipectinate (
umbrosa Hampson and viridumbrosa sp. n.). The venation is of the groundplan type. The forewings have a cryptic, variegated pattern in shades of brown or green, paler components of the irregular fasciae often being conspicuous. The hindwings are usually a uniform dark brown.

The male abdomen has the seventh segment elongated relative to the rest, and the sclerites of the eighth segment are modified, their basal margins cleft and bilobed. The genitalia are highly distinctive, but remarkably uniform through the genus, specific differences being minor and mainly in the aedeagus vesica. The uncus is short, rather rectangular, the gnathal arms leading into a thread-like subscaphium. The tegumen is elongate on each side, with a slight central flexure and a zone of large setal scars in the section ventral to the flexure. The transtilla and saccular shield are rather elongate, the valves arising from a narrow neck, paddle-like but folded so the ventral margin, with its numerous rows of small peg-like structures, overlaps the costal zone including the subbasal process. There are long setae on broad scars just distal to the subbasal process (Fig 429). There is a short saccus, stepped in on each side to produce a narrower apical component. The aedeagus is long, narrow with a spatulate basal part and a vesica with small cornuti.

 


In the female genitalia the lamella antevaginalis is developed as a setose pad, often diamond-shaped and distally bilobed or cleft, the ductus is short, the bursa small, rather rectangular, with a cluster of moderate spines at its centre, directed basad; such a cluster is present in some, but not all, species as mentioned later.

The greatest diversity is Oriental, possibly in Borneo where further species may await description and discovery as discussed below, but the genus attenuates eastwards to the Solomons and Queensland.

Bell (MS) reared species in India such as
semipars Walker. The larva is light grey with black transverse bands, and strongly suffused yellow. The head is yellow, somewhat heart-shaped. The body is slender, cylindrical, only slightly broader centrally. The prolegs are all present. Primary setae only occur. Gardner (1948a) also reared semipars and gave details of the chaetotaxy.

The larvae live stretched out on the undersides of the young leaves that they eat. They are somewhat sluggish. Pupation is on the underside of a leaf or elsewhere, in a slender, boat-shaped cocoon, with tapering processes extending fore and aft from the extremities of the ‘deck’, with one end of the ‘hull’ more or less vertical, the other curving convexly under like the bow of a boat. Viewed from above, the cocoon is lenticular. The cocoon is of densely woven, light orange silk, but the processes are medium brown or purplish, with this colour extending round the ‘deck’ as a dark line. The pupa is broadly rounded anteriorly, three times as long as broad, the posterior end also broadly rounded, lacking hooklets but beaded with longitudinal ridges along the anterior margin of the terminal segment. No rustling was noted for pupae in the genus, despite this beading.

The host plant was Diospyros (Ebenaceae), as it was also for L.
achine Felder, another species reared by Bell. Gardner (1948a) reared semipars from Glochidion (Euphorbiaceae).

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