This subfamily is revived, as indicated on (see
Introduction), to contain those
delicately built "oenochromine" genera with elongated, slender appendages
that tend to frequent the understorey of the tropical forests where they occur,
resting on the undersides of leaves by day, whence they can often be disturbed
into flight. There are no unambiguous synapomorphies to unite the two included
tribes, the Desmobathrini and Eumeleini: the modification of the tegumen of the
male genitalia occurs in a different manner in each; the reduction of cremastral
spines from eight to four, set on a disc or semicircular plate, is currently
known from only one of the Desmobathrini genera as well as Eumelea, so
cannot be considered a general feature of the group. However, each tribe can be
defined on much stronger characteristics.
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