TRIBE BOARMIINI
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Acrodontis Wehrli

Type species: aenigma Prout, Taiwan.

Though the facies is atypical of the Boarmiini, the presence of a cucullus on the valve and the general structure of the male genitalia, indicate this genus belongs within the boarmiine complex. The pupal cremaster in the two Japanese species (material sent to the author by Dr. R. Sato) is robust, rather asymmetric, bearing at least four major spines. This may indicate that the concept of the Boarmiini presented here is paraphyletic, with at least Acrodontis branching off more basally from most of the Boarmiini than do the three other tribes with a bifid cremaster.

The insect is large with an irregularly falcate forewing apex. The pattern consists of irregular mottling and blotching of rufous, pink and grey on a pale yellow ground; darker post medial fasciae are present, more or less punctate, and a broad, diffuse, discal spot on the forewing. The pattern is repeated on the underside.

The male antennae are bipectinate almost to the apex, those of the female filiform. In the abdomen, the tympanic bullae are relatively small. A fovea is present on the male forewing.

In the male genitalia the gnathus is robust, the tongue-like apical portion coarsely scobinate. The cucullus of the valve is well developed, the zone with coarse, short setae extending well basal along the costal margin and having a central lobe in the type species and the Japanese A. fumosa Prout. The sacculus terminates distally in a quadrate process, and a band of sclerotisation extends from it towards the costa. The aedeagus vesica bears scobination, often coarse, but no major cornuti; the apex of the aedeagus extends as sclerotised bands that bear spines, one or several.

The female genitalia have an extensively sclerotised lamella postvaginalis, with a system of fine folds surrounding a central zone that is less scobinate than the rest. The ductus is short, the base of the tubular section of the bursa scobinate and irregularly folded transversely. In the pyriform distal portion there is a transverse, mouth-shaped signum, a modification of the usual ‘mushroom’ type.

Larvae of the two Japanese species were illustrated by Sugi (1987). They are robust, finely striped longitudinally with white and blue-black, with irregular red or yellow patches along the flanks in association with the legs, prolegs and spiracles. One species is polyphagous but A. fumosa is restricted to Stachyurus (Stachyuraceae). This monobasic plant family extends from the Himalayas to Japan but is not known from Borneo or eastwards.

The genus is restricted to China, Japan and Korea apart from the new species described below.

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