Niaccaba Walker
Type species: sumptualis Walker, see below.
This is a
monobasic genus with facies as described for the type species below. Its
placement in association with the Araeopteroninae is contrary to the
arrangement of Hampson (1910), who associated the genus with Eublemminae, and
needs further investigation. No unambiguous pointers were identified in the
abdominal features noted below.
Features of the
head and wing venation were illustrated by Hampson (1910). The labial palps are
directed forwards, slightly upcurved, about twice the length of the head, with
the third segment very short. The male antennae are ciliate. The radial sector
veins are reduced, with R1 arising independently from the cell. The next two
radial veins have a common stalk, with the posterior branch reaching the apex
of the wing; the fourth is connate with the stalk of this pair. M3 and CuA1 are
stalked on both fore- and hindwings. There are no obvious phragma lobes between
the first and second abdominal tergites.
The male abdomen
has an eighth tergite as in the framed corematous condition, with apodemes
converging on a central thickening, but the sternite is three times as wide,
without a framed structure, except for a small central apodeme with narrow
lateral ridges running from it on the anterior margin. The genitalia have a
highly complex structure to the tegumen, juxta and vinculum as illustrated;
this is possibly a further modification of the complex Araeopteroninae
situation. The uncus is irregularly shaped, deeper over the basal half, with a
slight ‘ball and claw’ to its apex. The valves are narrow with a slender
saccular process interiorly.
The female
genitalia have the apodemes of the eighth segment reduced, but there are slight
lobes laterally on the anterior margin. The ductus is unsclerotised (therefore
not typically eublemmine) and extends for the length of the seventh segment.
The corpus bursae is narrow, rather sausage‑like, but with a hooked
appendix bursae at its basal end giving rise to the ductus seminalis. Distal to
this are some complex sclerites with a slender spine extending distad from them
amid some scobination. The distal half of the corpus bursae is unornamented.
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