Amyna Guenée
Type species: selenampha Guenée = punctum Fabricius, see below.
Synonyms: Amynodes Warren (type species distigmata Hampson, Bhutan); Berresa Walker (type species natalis Walker, see below); Chytoryza Grote (type species tecta Grote = axis Guenée, see below); Formosamyna Strand (type species frontalis Strand, Taiwan); Hesperimorpha Saalmüller (type species paradoxa Saalmüller, Madagascar = punctum); Hurworthia Nye (replacement name for Niphosticta Turner);
Ilattia Walker (type species cephusalis Walker = axis, see below); Lochia Walker (type species apicalis Walker, Queensland); Niphosticta Turner (type species apicipuncta Turner, Queensland), praeocc.; Pteraetholix Grote (type species bullula Grote, Alabama); Stridova Walker (type species albigutta Walker = axis, see below); Trilophia Turner (type species niphadospila Turner = auriculata Turner, Australia).
All species have
rather triangular forewings in shades of brown or blackish brown. The reniform
stigma may be prominent in the shape of a figure 8 and is sometimes variably
bounded by white and centred with white, pale brown or ochreous yellow; the
posterior ellipse or circle of the reniform may be more developed in this manner
than the anterior one. There are finely darker, crenate fasciae crossing the
wing but these tend to be obscure. In some species these are marked by whitish
marks at the costa, particularly the submarginal. Males of most species have a
distinctive fovea situated subbasally within the cell; this fovea is most
clearly seen on the underside, and may be partially covered by a flap of
scales. It is most strongly developed in natalis, and has been shown in
that species to have a stridulatory function (see p. 42). The labial palps are
porrect, the third segment half the length and depth of the second. The male
antennae are filiform, with short cilia. The phragma lobes between the first
and second abdominal tergites are vestigial.
The male abdomen
has the eighth tergite paddle‑like as in most Bagisarinae, though this
ranges from very narrow, almost violin‑shaped in punctum to very
broad (aroa Bethune‑Baker group). The sternite is more or less
uniformly sclerotised in punctum but has a framed lacuna in most of the aroa
group. The genitalia are characterised in particular by a scaled, shouldered
tegumen and by valves divided into a sclerotised costal process and a longer
narrower cucullar one that bears fine setae (seen also on the costal one in punctum).
There is usually a basally directed, curved or digitate process arising
subbasally from the central line of fusion of the two parts. The valve bases
are complex, but there is some indication of fusion of membranous extensions
from them basal to the juxta. The aedeagus is small, short, with a small,
irregularly globular vesica that may bear small groups of spines of varying
length or be immaculate.
The female
genitalia are variable, but the ostial area is unmodified, the ductus bursae
varies from moderate, sclerotised in punctum to short, unsclerotised
(e.g. all other Bornean species). The ostium is situated at the anterior margin
of the eighth segment. The corpus bursae is usually ovate to pyriform, small,
flimsy, without signa, but in punctum it is very narrow, equal in width
to the ductus.
The larvae have a
granulate head, and the prolegs of A3 and A4 are reduced to minute pointed
tubercles; the relatively long crochets of the developed prolegs are
appendiculate as in the Bagisarinae (p. 23) and typical members of the genus Cosmophila Boisduval (Gardner, 1941, 1946a, 1947; Holloway,
1998, 2005). The body is slender, nearly cylindrical. Features of the pupa are
also bagisarine (p. 25).
The genus is
pantropical. Indo‑Australian species from Sundaland eastwards are
reviewed here in some detail. In addition to those discussed in the species
accounts, there are three further species in Australia (Nielsen et al.,
1996): auriculata Turner and onthodes Lower, with more variegated grey forewings; apicalis Walker, with rather narrow forewings but male
genitalia similar to those of axis.
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