TRIBE BOLETOBIINI
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Panilla Moore

Type species: dispila Walker, type locality not known [India, Sri Lanka, Nepal].

Most species have facies like the Bornean ones, resembling somewhat small boarmiine geometrids, but usually with darker brown triangles or other patches on the forewing costa and obliquely posterior to it, both immediately distad to the postmedial. The third segment of the labial palp is short.

The male abdomen has the eighth segment typical of this fungus-feeding group. The genitalia may be tufted with scales (e.g. particularly
combusta Hampson; see below). The uncus is relatively long and slender. The tegumen has lobes centrally on each side and there is a slender looped structure from the base of the valve costa that curves up inside the tegumen within the diaphragma. The juxta is a catocaline inverted ‘V’. The valves terminate in a complex set of two or three processes of diverse shape. The aedeagus vesica is more globular than in related genera and scobinate or with a few small spines.

The female (
terminalis Hampson comb. n.; see below) is similar to that of Caduca Walker (see below) in having a central band of spicules (spicules similar to those in Artigisa) in the corpus bursae, but the appendix bursae is associated with this zone of spicules and thus in a more distal position. In homospila Hampson, the appendix bursae arises more basally and the spining is more distal. The ostium is between the seventh and eighth segments as in Caduca and Artigisa rather than obliquely within the eighth as in Drepanorhina and Diomea Walker.

Bell (MS) described the larva of the type species in S. India. Prolegs are absent from A3 and A4. The head is round, with a well developed vertical sinus dividing it into two broadly rounded lobes; its surface is rough, dull, light yellowish, marbled with black, with little tubercles, and setae are present. The body is dull, with primary setae on prominent, rather cylindrical tubercles. These setae are black or white, sinuous, with some even plumose. The segments are each ochreous brown but mostly ringed black anteriorly at the junction with the next. There is dull yellow marbling and the hint of orange subdorsal and lateral bands that are interrupted at the segmental junctions.

The larva lives on fungus, often colonially, walking with a strong looping motion. Pupation is in an ovoid cocoon incorporating particles of frass and other substrate.

The genus as currently recognised (Poole, 1989) extends through the Indo-Australian tropics to Australia and the Solomons, and there are also four species in Africa. Yoshimoto (2001b) has reviewed and illustrated many of the mainland Asian species.

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