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Nola
Leach
Type
species: palliola
[Denis Schiffermüller] (=
cucullatella
Linnaeus), Austria.
Synonyms: Aradrapha
Walker (type species
partitalis
Walker, S. Africa);
Argyrophyes
Grote (type species
cilicoides
Grote, U.S.A.);
Automala
Walker (type species
semidolosa
Walker, Borneo);
Celama
Walker (type species
liparisalis
Walker = bifascialis Walker, Borneo);
Celamoides
van Eecke (type species
pseudastigma
van Eecke, Java) syn. n.;
Chlamifera
Hübner (type species palliola [Denis & Schiffermüller]);
Deltapterum
Hampson (type species
peguense
Hampson, Burma);
Epizeuctis
Meyrick (type species
innocua
Butler, Taiwan);
Idiocyttara
Turner (type species
tornotis
Meyrick, Australia);
Lebena
Walker (type species
trinotata
Walker = triquetrana
Fitch, N. America);
Lira
Billberg (unnecessary replacement name for Nola);
Minnagara
Walker (type species
fasciata
Walker, Moluccas);
Necla
Walker (type species canioralis Walker, Borneo);
Neonola
Hampson (type species mesosticta Hampson, Borneo);
Pisara
Walker (type species opalina Walker, Borneo);
Roeselia
Hübner (type species
cucullatella
Linnaeus, Europe).
Species in this genus typically have reduced forewing and hindwing venation as
in Manoba,
though in some taxa (e.g. the type species of
Pisara
and Neonola)
the radial sector venation is further reduced to R1 and a bifurcation
representing the rest. The male antennae are bipectinate or fasciculate, the
latter condition occurring in most Bornean species.
The genus is most clearly defined on features of the male genitalia
(Franclemont, 1960; Holloway & Miller, 1995): loss or extreme reduction of the
uncus; absence of scaphial structures; cleavage of the valves into two
longitudinal sections, the more ventral one bearing the harpe but also often
having a small spine near its apex (e.g. Fig 108). The apodemes of the eighth
segment are present on the tergite and often also the sternite, and usually more
separated than in Manoba.
The female genitalia are typical of more advanced members of the subfamily, with
two horn-like invaginations forming the signa of the corpus bursae, sometimes
reduced to one or to adjacent scobinate domes (Figs 96, 97).
The genus is extremely widespread and diverse, but its true extent will only be
gauged when taxa currently assigned to it have this placement supported by
dissection of the male genitalia. Host records (Robinson et al., 2001)
include several instances of feeding on flowers, fruits and pods; one instance
of gall-feeding has been noted (Itō & Hattori, 1982).
Several further genus-group names placed as synonyms of Nola by Poole
(1989) and Nielsen et al. have either been referred to under
Meganola or Manoba or have not been checked in the light of the
more strict definition above. These latter are:
Cryphimaea
Turner (type species
poliophasma
Turner, Queensland, Australia);
Neocleta
Turner (type species
empyra
Turner, Western Australia);
Nestiodes
Turner (type species
eremnopa
Turner, New South Wales, Australia);
Stenola
Möschler (type species
bistriga
Möschler, Puerto Rico).
Poliothripa
Hampson Gen. rev. (type species
niphostema
Hampson (praeocc.) =
steniphona
van Son, S. Africa) should also be excluded; the genitalia of both sexes are
distinct from those of Manoba.
A
few trifine species that do not fall readily into Nola,
Manoba
or these other genera are treated under
"Nola”.
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