TRIBE NUDARIINI
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Cyana costifimbria Walker  
   
Bizone costifimbria Walker, 1862, J. Linn. Soc. (Zool.), 6:121.
   
Chionaema rhadota Swinhoe, 1907, Ann. Mag nat. Hist. (7), 20: 76, syn. n.
   
Chionaema javanica ab. niasensis Strand, 1922, Lep. Cat., 26: 685, syn. n.
   
Chionaema rhadota borneensis Talbot, 1926, Sarawak Mus. J., 3(2): 135, syn. n.
   
Cyana pectinata Roepke, [1946a]: Tijdschr. Ent. 87: 31, syn. n.


Cyana costifimbria
(approx. lifesize)

Cyana costifimbria
(approx. lifesize)


Diagnosis.
This species shows extreme sexual dimorphism. Males are unmistakeable, with transverse broad orange bands on the forewing (very rarely edged with some black) being disrupted by a long and conspicuous fringe of scales extending back from the costa. Females are like a small version of those of conclusa or malayensis, but have the postmedial black edge mostly straight, but angled obtusely towards the apex at the costal end. The antemedial black edge meets the dorsum at an acute angle basad as in malayensis rather than at an obtuse one as in conclusa.

Taxonomic note. The marked sexual dimorphism has led to some confusion in the past in that both sexes have been described more than once and rarely associated with each other (see also C. determinata Walker). A related species is C. vespertata Cernư comb. n. from the Philippines; this shows similar sexual dimorphism, but the male has black delineation of the fasciae. The group is distinguished by a spur on the centre of the valve costa, a ridge between the two valve sections, and general scobination of the aedeagus vesica, including two patches of longer spines.

Geographical range. Borneo, Peninsular Malaysia, Sumatra, Nias, Java, Philippines (Palawan, Tawi Tawi).

Habitat preference. The species has a similar ecological range to determinata (see Cyana determinata Walker) but is slightly commoner.

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