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Comibaena Hübner

Type species: bajularia Denis & Schiffermüller = pustulata Hufnagel, Europe.

Synonyms: Chlorochaeta Warren (type species longipennis Warren, Nigeria); Colutoceras Warren (type species diluta Warren, Japan); Phorodesma Boisduval (type species bajularia); Probolosceles Warren (type species biplaga Walker); Uliocnemis Warren (type species cassidara Guenée) syn. n.

This and the next three genera represent the Comibaeniti in Borneo, and share the general characters noted on (Geometrini). The male antennae are strongly bipectinate.

Species of Comibaena have brown patches at the forewing tornus and hindwing apex as in Rhomboristiti, though in some, including the type species, these are associated with submarginal and marginal white markings, a feature in common with Protuliocnemis Gen. n. below. There is usually a black discal spot on the hindwing.

The best diagnostic features for Comibaeniti genera appear to be in the male and female genitalia. Comibaena is here characterised by the presence of a bifid uncus with strong socii, giving a quadrifid uncal structure, a weak, ribbon-like gnathus, a variably sclerotised and ornamented valve costa (seen in other Comibaeniti genera), and modification in many cases to the eighth sternite. In the female there is usually a sclerotised postvaginal projection.

The larva is generally shaped and behaves as described below. Specific accounts for C. cassidara and species related to C. biplaga Walker are presented below. Japanese species are recorded as feeding on numerous plant families (Nakajima & Sato, 1979), and several larvae are illustrated in Sugi (1987).

The genus is found widely in the Old World, including temperate latitudes, but is perhaps most diverse in the Oriental Region.

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