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This checklist includes all known Bomean species, giving the
page number on which a fuller account can be found, a broad categorisation of
the type of geographical distribution and Bornean habitat preference.
In the geographical distribution the term Wallacea indicates
the presence of the species in Sulawesi and the Philippines, *
means shared with
Sumatra only and
**
means shared with Peninsular Malaysia only. The suggested
habitat preference is bracketed if based on less than five specimens. The
categories 'Radio Sabah' and 'Summit' refer to zones on G. Kinabalu above upper
montane recognised by Holloway (1970).
OENOCHROMINAE (6 species)
Sarcinodes
vultuaria
Guenée. Thailand, Sundaland, Sulawesi. Lowland forest.
Sarcinodes reductatus Inoue.
Sundaland. Lower and upper montane
Sarcinodes tornubilatus
Sommerer. Sundaland. Lower and upper montane.
Sarcinodes malakarius
Sommerer.
Sundaland. (Lower and upper montane).
Sarcinodes sumatraria Walker.
Sundaland. (Lowland forest).
Sarcinodes perakaria
Swinhoe. Sundaland. (Lowland to
upper montane forests).
DESMOBATHRINAE: DESMOBATHRINI (35 species)
Ozola
edui
Sommerer. Sundaland. (Lowland).
Ozola
basisparsata Walker. Borneo, Engano, Philippines, (?Lowland)
Ozola
turlini
Herbulot. Sundaland, Sulawesi. Lowland.
Ozola
hollowayi Scoble
& Sommerer. Sundaland to Bismarcks, Queensland. No habitat data.
Ozola
minor Moore. Oriental tropics. Lowland.
Ozola
impedita Walker. Endemic. (Lowland to lower
montane).
Ozola
pannosa Holloway. Sundaland*. Lowland forest (to upper montane).
Ozola
falcipennis Moore. N.E. Himalaya to Sundaland**. Lower and upper montane.
Ozola
liwana Sommerer. Sundaland*. Upper montane.
Ozola
apparata Prout. Sundaland. (Upper
montane).
Ozola
prouti Holloway. Endemic. Upper
montane.
Ozola
submontana Holloway. Endemic. Lower and upper
montane.
Noreia
ajaia Walker. Oriental tropics to Sundaland. Lowland forest, (lower
montane).
Noreia
unilineata Walker.
Sundaland. Lowland forest.
Noreia
achloraria Warren.
Sundaland, Sulawesi. Lowland forest.
Noreia
anacardium sp.
n. Endemic. (Lowland forest), lower (and upper) montane.
Noreia
sinuilineata sp.
n. Endemic. (Lowland forest).
Foveabathra
venusta Warren.
Sundaland. (Lowland).
Conolophia
nigripuncta Hampson. Indian Subregion, mainland
S.E. Asia, Borneo. (Lowland), lower and
upper montane.
Alex
palparia Walker. Himalaya to Sundaland, Philippines, Lesser Sundas. Lowland and lower
montane forests, (upper montane).
Celerena
signata Warren.
Sundaland. Lowland, particularly disturbed forest.
Derambila
saponaria Guenée.
Indian Subregion, Sundaland. No habitat data.
Derambila
fragilis Butler. Oriental tropics. (Lowland).
Derambila
zanclopterata Walker.
Sundaland. (Lowland).
Derambila
costata Warren. N.E. Himalaya,
Sundaland*. (Lowland).
Derambila zincaria Guenée. Sundaland to New Guinea. Lowland
disturbed and open habitats.
Derambila dentifera Moore. Oriental tropics to Sundaland,
Philippines. (Lowland plantations).
Derambila propages Prout. Endemic. (Lowland).
Derambila dentiscripta Bastelberger. Sundaland*, Sulawesi,
Moluccas. (Lowland, upper montane).
Derambila lumenaria Geyer. Indian Subregion to Sundaland.
(Lowland).
Derambila herbuloti sp. n. Endemic. (Lowland).
Derambila gertraudae Sommerer. N.E. Himalaya, Sundaland*.
(?Lower montane).
Derambila manfredi sp. n. Endemic. (Lower montane forest).
Derambila manca Swinhoe. Endemic, N. Sarawak. Lowland alluvial
forest.
Derambila livens Prout. Endemic. (Lowland forest).
DESMOBATHRINAE:
EUMELEINI (10 species)
Eumelea
rosalia Stoll. Indo-Australian
tropics. (Lowland).
Eumelea feliciata Guenée. N.E. Himalaya, Sundaland*. (Montane).
Eumelea unilineata Warren. Borneo, Philippines. (Lowland).
Eumelea florinata Guenée. N.E. Himalaya to Sundaland. Lowland
forest.
Eumelea biflavata Warren. N.E. Himalaya to Sundaland. Lowland
forest.
Eumelea ludovicata Guenée. Indo-Australian tropics. No habitat
data.
Eumelea rubrifusa Warren. Borneo, Wallacea. Upper
montane.
Eumelea parda Sommerer. Sundaland. (Lowland)
Eumelea djingga Sommerer?. Sundaland*. (Lower montane).
Eumelea obesata Felder & Rogenhofer. Borneo, Philippines.
Upper montane.
GEOMETRINAE:
DYSPHANIINI (8 species)
Dysphania
militaris Linnaeus. Oriental
Region to Sundaland. (?Lowland).
Dysphania sagana Druce. S.E. Asia, Sundaland. (Lowland).
Dysphania subrepleta Walker. N.E. Himalaya to Hainan and
Sundaland. Lowland to upper montane.
Dysphania bivexillata Prout.
Sundaland. Mangrove.
Dysphania discalis Walker. Sundaland**. (Montane).
Dysphania malayanus Guérin-Méneville.
Sundaland. Lowland
forest.
Dysphania transducta Walker. Burma, Sundaland. Lowland forest.
Dysphania glaucescens Walker.
Sundaland. (Lowland).
GEOMETRINAE:
GEOMETRINI (209 species)
Orthorisma netunaria Guenée. Sundaland*. (Lowland, lower montane forest).
Actenochroma muscicoloraria Walker. N.E. Himalaya, Sundaland*.
(Lowland).
Herochroma urapteraria Walker.
Sundaland. (Lowland), lower and
upper montane.
Herochroma subtepens Walker. Sundaland. (Lowland, lower montane
forests).
Herochroma orientalis Holloway.
Indian Subregion, Sundaland.
(Lower and upper montane).
Herochroma xuthopletes Prout. Sundaland*. (Lowland forest).
Herochroma flavibasalis Warren. Sundaland. (Lowland) and lower montane forest.
Herochroma clariscripta sp. n. Endemic. (Lowland).
Sundadoxa multidentata Prout. Sundaland. Lowland and lower
montane forest.
Pullichroma pullicosta Prout. Sundaland*. Wallacea. (Upper
montane forest).
Metallolophia vitticosta Walker. Sundaland. Lowland (and lower
montane) forest.
Metallolophia subradiata Warren. Sundaland. (Lowland) and lower
montane forest.
Metallolophia variegata
sp. n. Endemic. (Lowland) and lower
montane forest.
Metallolophia cineracea sp. n.
Sundaland**. (Lowland heath
forest).
Epipristis truncataria Walker. Sundaland. Lowland.
Epipristis nelearia Guenée. Indo-Australian tropics to
Queensland. Lowland.
Pingasa ruginaria Guenée. Indian Subregion to Sundaland.
Lowland (to montane).
Pingasa chlora Stoll. Sundaland, Wallacea, Moluccas, Queensland.
Disturbed and cultivated areas, lowland to lower montane.
Pingasa lariaria Walker. Indo-Australian tropics from N. India
to New Guinea. Lowland to upper montane.
Pingasa rubicunda Warren. N. India, Sundaland, Philippines.
Lowland forests, including disturbed areas and plantations, (lower montane
forest).
Pingasa rubimontana Holloway & Sommerer. Sundaland*,
Sulawesi. Upper montane.
Pingasa tapungkanana Strand. Sundaland. Lowland to upper
montane.
Pingasa subviridis Warren. India, Sundaland. Lowland to upper
montane forests.
Pingasa subpurpurea Warren. Borneo, Wallacea. (Lowland).
Pingasa venusta Warren. N.E. Himalaya to New Guinea. Lowland to
lower montane forest, often in areas with disturbed or secondary forest.
Pachyodes pratti Prout. Sundaland. (Lowland and lower
montane).
Lophophelma vigens Butler. Himalaya, Sundaland. Lower and upper
montane.
Lophophelma rubroviridata Warren. Sundaland. Lower and upper
montane.
Lophophelma luteipes Felder & Rogenhofer. N.E. Himalaya,
Sundaland. Lower and upper montane.
Lophophelma funebrosa Warren. N.E. Himalaya, Sundaland, Lesser
Sundas. (Lowland).
Lophophelma loncheres Prout. Sundaland. Lowland forest.
Lophophelma erionoma Swinhoe. N.E. Himalaya, W. China, Sundaland.
Lower and upper montane.
Dindica alaopis Prout. Endemic. Lower and upper montane.
Dindica polyphaenaria Guenée. Himalaya, SE. Asia, Sundaland.
(Lowland).
Dindica olivacea Inoue. N.E. Himalaya to Borneo, Sumatra,
Philippines. Lowland.
Tanaorhinus malayanus Inoue. Sundaland. (Upper montane).
Tanaorhinus rafflesii Moore. Sundaland. Lowland to upper montane.
Tanaorhinus viridiluteata Walker. N.E. Himalaya to Taiwan and
Sundaland. Lowland to upper montane.
Mixochlora vittata Moore. Himalaya to Japan and Sundaland.
(Lower) to upper montane.
Mixochlora argentifusa Walker. Sundaland, Sulawesi. (Lowland
forest).
Timandromorpha energes Prout. Thailand, Sundaland. Lower to
upper montane.
Chloroglyphica xeromeris Prout. Sundaland. (Lowland), upper
montane.
Paramaxates polygrapharia Walker. Sundaland, Sulawesi. Lowland
forest, (lower and upper montane).
Paramaxates posterecta Holloway. N.E. Himalaya to Taiwan and
Sundaland. (Lowland). Upper montane forest.
Paramaxates macrocerata Yazaki. Sundaland. (Lowland).
Paramaxates yazakii sp. n. Sundaland. Lower and upper montane.
Paramaxates spinivesica sp.
n. Endemic. No habitat data.
Dooabia
plana Prout. Endemic. Upper montane.
Dooabia
lunifera Moore. N.E. Himalaya, Sundaland*. Upper montane.
Dooabia
puncticostata Prout
. Sundaland. Lowland (to montane).
Dooabia
bruneiconcha sp.
n. Endemic. (Lowland forest).
Euxena
albiguttata Warren.
Sundaland, Sulawesi. Lowland.
Chlorodontopera
chalybeata Moore. N.E. Himalaya, mainland S.E. Asia, Sundaland*. (Lowland).
Agathia
succedanea Warren. Sundaland*. Lower and upper montane.
Agathia
laetata Fabricius. Indian Subregion to Taiwan and Sundaland. Lowland forest.
Agathia
quinaria Moore. N.E. Himalaya, Sundaland. Lowland (and lower montane) forest.
Agathia
largita sp.
n. Sundaland*. (Lowland).
Agathia
arcuata Moore. Indian Subregion to Hainan and Sundaland. Lowland forest.
Agathia
deliciosa Holloway
& Sommerer. Sundaland*. (Lowland forest).
Agathia
rubrilineata Warren.
Sundaland. (Lowland and lower montane forest).
Agathia
eromenoides sp.
n. Sundaland**. (Lowland forest).
Agathia
diplochorda Prout
. Sundaland. (Lowland and lower montane forest).,
Agathia
codina Swinhoe. N.E. Himalaya, Sundaland. (Lowland forest).
Agathia
obsoleta Warren.
Sundaland, Philippines. (Lowland forest).
Agathia
gigantea Butler.
Sundaland. (Lowland forest).
Agathia
vicina Bastelberger.
Sundaland. (Lowland forest).
Agathia
cristifera Walker
. Sundaland. (Lowland forest).
Agathia
muluensis sp.
n. Endemic. Lower montane forest.
Agathia
laqueifera Prout. N.E.
Himalaya, Sundaland. Lowland forest.
Agathia
spp. Possibly two further lowland species of the cristifera group occur
in Borneo.
Ornithospila
avicularia Guenée. N.E. Himalaya, Sundaland*, Philippines. Lowland forest (and lower
montane).
Ornithospila
bipunctata Prout
. Sundaland, Wallacea. Lowland to upper montane.
Ornithospila
submonstrans Walker.
Sundaland, Philippines, Moluccas. Lowland to upper montane.
Ornithospila
sundaensis Holloway.
Sundaland. Lowland and lower montane forest, (upper montane).
Ornithospila
cincta Walker.
Sundaland. Lowland forest, (montane).
Ornithospila
succincta Prout.
Sundaland, Philippines, New Guinea. Lowland to upper montane.
Ornithospila
lineata Moore. Indian Subregion, mainland
S.E. Asia, Sundaland*. (Lowland, upper
montane).
Ornithospila
esmeralda Hampson. N.E. Himalaya, Sundaland, Philippines. (Lowland, upper
montane).
Eucyclodes
gavissima Walker. Indian Subregion to Taiwan and Sundaland*. (Lowland forest).
Eucyclodes
charma Prout.
Sundaland. (Lowland).
Eucyclodes
textiloides sp.
n. Sundaland. (Lowland forest).
Eucyclodes
albisparsa Walker. Indian Subregion, Sundaland, Sulawesi. (Lowland) and lower montane
forest.
Eucyclodes
rufimargo Warren. Sundaland**, Sulawesi. (Lowland forest).
Eucyclodes
discipennata Walker.
Sundaland. (Lowland to upper montane).
Eucyclodes
semialba Walker. Indian Subregion to Sundaland. Lowland forest, (upper
montane).
Eucyclodes
discata Warren. Endemic. (Lowland, lower and) upper
montane.
Rhombocentra
semipurpurea Warren. N.E. Himalaya, Sundaland*. Lowland forest.
Spaniocentra
megaspilaria Guenée. Endemic. Lowland forest (and lower
montane).
Spaniocentra
apatelloides sp.
n. Endemic. Upper montane.
Spaniocentra
lobata Holloway. Endemic. Lowland forest (and lower
montane).
Comibaena
albimarginata Warren. N.E. Himalaya, Sundaland*. Upper montane.
Comibaena
biplaga Walker. N.E. Himalaya, Sundaland. Lowland forest (and lower
montane).
Comibaena
cassidara Guenée. India to China and Borneo. (Cultivated areas in lowland and lower
montane zones).
Comibaena
fuscidorsata Prout. N.E. Himalaya, Sundaland*. Lowland forest (and lower
montane).
Comibaena
attenuata Warren.
Sundaland, Wallacea. Lowland forests including plantations.
Protuliocnemis
partita Walker. Indo-Australian tropics. Lowland forest, (upper
montane).
Protuliocnemis
helpsi sp.
n. Borneo, Singapore. Mangrove.
Protuliocnemis
biplagiata Moore. Indo-Australian tropics. Lowland (to upper
montane).
Comostolodes
albicatena Warren. N.E. Himalaya, Sundaland*. (Lower and upper montane).
Comostolodes
dialitha West. Sundaland*, Java, Philippines. Lowland forest (and lower montane).
Argyrocosma
strepens Prout. Sundaland*, Sulawesi. (Lowland).
Argyrocosma
inductaria Guenée. Indian Subregion, Borneo. Lowland forest.
Argyrocosma
consobrina Warren. Borneo, Java to Bismarcks and Queensland. (Lowland forest).
Aporandria
specularia Guenée. Oriental tropics. Lowland, including disturbed areas.
Oenospila
flavifusata Walker. Oriental tropics to Sundaland. Lowland forests and plantations.
Oenospila
microstrix sp.
n. Sundaland*. (Lowland forest).
Oenospila
altistrix
sp. n. Endemic. (Lower and upper montane).
Olerospila
oleraria Guenée. Sundaland to Solomons. (Lowland forest,
montane).
Episothalma
robustaria Guenée. Indian Subregion to Sundaland. (Lowland forest).
Eretmopus
marinaria Guenée. Thailand to New Guinea. Mangrove.
Thalassodes
opalinoides sp.
n. Endemic. Lowland to upper montane.
Thalassodes
depulsata Walker. Borneo, Sulawesi. (Lowland alluvial forest).
Thalassodes
immissaria Walker. ?Oriental tropics. (Lowland).
Thalassodes
sundissepta sp.
n. Sundaland. Lowland forest, (lower montane).
Thalassodes
dissitoides sp.
n. Andamans, Sundaland, Wallacea. Lowland forest. (lower montane).
Thalassodes
linguissita sp.
n. Sundaland*. (Lowland forest).
Thalassodes
viridifascia Swinhoe. Borneo, Sulawesi, Seram. (Lower
montane).
Thalassodes
curiosa Swinhoe. N.E. Himalaya, Sundaland, Sulawesi, S. Moluccas. Upper
montane.
Orothalassodes
hypocrites Prout. N.E. Himalaya, Sundaland**. (Lower montane).
Orothalassodes
retaka sp. n. Endemic. (Upper
montane).
Orothalassodes
leucoceraea Prout. Sundaland**. (Lowland heath forest, upper montane).
Orothalassodes
floccosa Prout.
Sundaland. (Lower and upper montane).
Orothalassodes
glabrosa sp.
n.. Endemic. (Lowland, upper montane).
Pelagodes
forceps sp.
n. Endemic. (Lowland, lower montane).
Pelagodes
cochlearis sp.
n. Sundaland*. (Lowland).
Pelagodes
rana sp. n. Endemic. No habitat data.
Pelagodes
waterstradti sp.
n. Endemic. (Montane).
Pelagodes
tridens sp.
n. Endemic. (Lowland).
Pelagodes
semengok sp.
n. Sundaland. (Lowland forest).
Pelagodes semihyalina Walker. Sundaland. Lowland forest.
Pelagodes spiniseparati sp. n. Sundaland**. (Lower montane
forest).
Pelagodes clarifimbria Prout. Sri Lanka, Sundaland**. (Lowland
forest).
Pelagodes falsaria Prout. India, Borneo, Java, Bali.
Upper montane.
Hemithea neptunaria sp. n. Endemic. Lower montane.
Hemithea krakenaria sp. n. Endemic. (Lowland forest).
Hemithea poseidonaria sp. n. Endemic. (Lowland) and lower
montane forest.
Hemithea undifera Walker. Sundaland. Lowland forest.
Hemithea insularia Guenée. Indo-Australian tropics. (Lowland
forest).
Hemithea antigrapha Prout. N.E. Himalaya, Sundaland**. (Lowland
and lower montane forest).
Hemithea marina Butler. Oriental tropics, Seram. Lowland forest.
Hemithea notospila Prout. Sundaland, Sulawesi. (Lowland, upper montane).
Hemithea melalopha Prout. N.E. Himalaya, Sundaland**,
Philippines, Seram. (Lowland forest).
Hemithea wuka Pagenstecher. Indo-Australian tropics. (Lowland Pinus
plantation).
Pamphlebia rubrolimbraria Guenée. Indo-Australian tropics
(Lowland).
Idiochlora pudentifimbria Prout. N.E. Himalaya, Sundaland**.
(Lowland).
Idiochlora celataria Walker . Borneo to New Guinea, Queensland.
(Lowland).
Idiochlora subexpressa Walker. Sundaland**? (Lowland).
Idiochlora innotata Walker. Sundaland*. (Lowland).
Idiochlora olivata Warren. Sundaland**. (Lowland).
Idiochlora berwicki Holloway. Endemic. (Upper montane).
Idiochlora stictogramma Prout. Endemic. (Upper montane).
Idiochlora stictogrammoides sp. n. Endemic. (Upper montane).
Albinospila floresaria Walker. Oriental tropics. Lowland forest,
(upper montane).
Maxates coelataria Walker. Indian Subregion to Sundaland.
(Lowland forest).
Maxates dysides Prout . Sundaland*. Lowland forest.
Maxates thetydaria Guenée. Indian Subregion to Taiwan, Borneo.
Lower and upper montane.
Maxates veninotata Warren. N.E. Himalaya, Borneo. (Lowland
forest).
Maxates waterstradti Prout. Endemic. (Lower and upper montane).
Maxates melinau sp. n. Endemic. (Lowland forest).
Maxates iridescoides sp. n. Endemic. (Lower montane).
Maxates magnipuncta Prout. Sundaland**. Lowland forest.
Maxates submontana sp. n. Endemic. Lower montane.
Maxates korintjiensis Prout
. Sundaland*. (Lower and upper montane).
Maxates melancholica Prout. Sundaland. (Lower
montane).
Maxates tristis sp. n. Endemic. (Lowland forest).
Maxates marculenta Prout. Endemic. Lower and upper
montane.
Maxates lugubriosa sp. n.. Borneo, Java. Lowland forest.
Maxates variegata sp. n. Endemic. (Upper
montane~.
Maxates obliterata sp. n. Endemic. Lowland forest.
Maxates muluensis sp. n. Endemic. Lowland forest.
Maxates seria sp. n. Sundaland*? (coastal/secondary forest).
Maxates sp. Endemic. (Lowland forest).
Chlorissa aquamarina Hampson. N.E. Himalaya, Borneo. (Lowland),
lower and (upper) montane forest.
Anoplosceles nigripunctata Warren. Sundaland. Lowland forest.
Chloristola setosa sp. n. Endemic. Lowland forest.
Jodis
nanda Walker. Oriental tropics. (Lowland).
Jodis
subtractata Walker. Oriental tropics to Sundaland. Lowland forest.
Jodis
spumifera Warren. Sundaland. (Lower and upper
montane).
Jodis
rectisecta Herbulot. Borneo, Palawan. (Lower
montane).
Jodis
api sp.
n. Endemic. (Lower montane).
Berta
vaga Walker. Sundaland*. (Lowland forest).
Berta
annulifera Warren. N.E. Himalaya to
Bismarcks. Lowland (and lower montane) forest.
Berta
chrysolineata Walker. Indo-Australian tropics. Lowland forest.
Berta digitijuxta sp.
n. Borneo, Java (Upper montane).
Berta
cercifera sp.
n. Borneo. (Lowland).
Berta
tridentijuxta sp.
n. Borneo, Sulawesi, Seram. (Lowland and lower montane forest).
Berta
albiplaga Warren. N.E. Himalaya to New Guinea. (Lowland forest).
Berta
anteplaga Prout. N.E. Himalaya, Borneo. (Lower
montane).
Berta
copiosa Prout. N.E. Himalaya, Sundaland. (Lowland forest, upper
montane).
Berta
zygophyxia Prout. Sundaland**. (Lower montane forest).
Berta
sp.. Sundaland*. (Lowland Pinus plantation).
Comostola
laesaria Walker. Indo-Australian tropics. Lowland forest.
Comostola
orestias Prout.
Sundaland, New Guinea. Lowland to upper montane.
Comostola
meritaria Walker. Sri Lanka, Taiwan, Sundaland*, Sulawesi. (Lowland and lower montane
forest.).
Comostola
nereidaria Snellen. Borneo, Sulawesi to Solomons. (Lowland forest).
Comostola
subtiliaria Bremer. N.E. Himalaya to Japan, Korea, Sundaland*. Upper
montane.
Comostola
cedilla Prout. Sundaland to New Guinea and Queensland. (Lowland forest).
Comostola
chlorargyra Walker. Oriental tropics. Lowland forest.
Comostola enodata sp. n. Walker. Sundaland, Sulawesi, New Guinea. Lowland
forest, including disturbed forest.
Comostola pyrrhogona Walker. Indo-Australian tropics. Lowland (and lower
montane) forest.
Comostola turgescens Prout. N.E. Himalaya, Sundaland, Sulawesi. (Lowland
and lower montane forest.
Comostola dyakaria Walker. N.E. Himalaya, Sundaland, Philippines. Lower montane.
Pseudocomostola cosmetocraspeda
Prout. N.E. Himalaya, Borneo. (Lowland and lower montane
forest).
Mystichlora mystica Prout. Sundaland*, Sulawesi. (Lowland forest).
Pseudidiochlora temburong sp. n. Endemic. (Lowland forest).
Rhanidopsis alleni sp. n. Sundaland*. (Lowland forest).
Geometrine
species A. Endemic. (Lowland heath forest).
Geometrine
species B. Endemic. (Lowland).
?ENNOMINAE
(1 species)
Heteralex rectilineata Guenée. Sundaland. Lowland and lower montane.
?ENNOMINAE:
ORTHOSTIXINI (2 species)
Naxa
guttulata Warren. Sundaland*. (Lowland, lower montane).
Naxa
kerangatis sp.
n. Endemic. Lowland heath forest.
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